Taxonomic Review of the Species Groups of the Genus Physalaemus Fitzinger, 1826 with Revalidation of the Genera Engystomops Jiménez-De-La-Espada, 1872 and Eupemphixsteindachner, 1863 (Amphibia, Anura, Leptodactylidae)
The species groups of the genus Physalaemus are redefined based on morphometrics, external morphology, color patterns, and osteological characters. Seven species groups of Physalaemus are recognized: P. cuvieri group (eight species), P. signifer group (ten species), P. albifrons group (four species), P. deimaticus group (three species), P. gracilis group (five species), P. henselii group (three species), and P. olfersii group (four species). The genus Engystomops Jiménez-de-la-Espada, 1872 is revalidated to include the former P. pustulosus group (sensu Lynch, 1970). It consists of seven species: E. petersi Jiménez-de-la-Espada, 1872 (type-species by monotypy), E. coloradorum (Cannatella & Duellman, 1984), E. guayaco (Ron, Coloma & Cannatella, 2005), E. montubio (Ron, Cannatella & Coloma, 2004), E. pustulatus (Shreve, 1941), E. pustulosus (Cope, 1864), and E. randi (Ron, Cannatella & Coloma, 2004). The genus Eupemphix Steindachner, 1863 is revalidated to include E. nattereri Steindachner, 1864.
